Châtelperronian and Uluzzian techno-complexes; Climate forcing and feedback; Last glacial period; Millenial-scale variability; Palaeotemperature proxies; Pollen assemblages; Western Europe climate; Chatelperronian; Chatelperronian and uluzzian techno-complex; Climate feedbacks; Climate forcings; Palaeotemperature proxy; Palaeotemperatures; Pollen assemblage; Western Europe; Western europe climate; Global and Planetary Change; Ecology, Evolution, Behavior and Systematics; Archeology (arts and humanities); Archeology; Geology
Abstract :
[en] Western European climate during the Last Glacial period (50–20 ka BP) was shaped by complex interactions between oceanic and terrestrial conditions, modulated by both orbital and millennial-scale forcings. This study integrates 75 marine and terrestrial palaeoenvironmental records to reconstruct spatiotemporal climate variability across Europe and the Mediterranean region. Sea surface temperature (SST) records reveal a pronounced latitudinal gradient, with strong cooling at high latitudes and relative SST stability at mid-latitudes on multi-millennial timescales, which likely moderated western European climate. Pollen-based vegetation records, in contrast, show a steep longitudinal trend, with intensified environmental shifts southeast of the Alps, where oceanic moderation was limited. On millennial timescales, Dansgaard-Oeschger cycles and Heinrich stadials imprinted distinct signatures on SST and vegetation, especially near the Atlantic margin, linked to disruptions in the Atlantic Meridional Overturning Circulation. These findings highlight the differentiated sensitivity of oceanic and terrestrial conditions to climate drivers and the climatic role of geographic barriers such as the Alps. The resulting spatial heterogeneity likely influenced human cultural trajectories, with stable long-term conditions but strong Heinrich stadial near the Atlantic coinciding with the emergence of the Châtelperronian industry, while more pronounced climatic long-term trend in southeastern Europe paralleled the development of the Uluzzian complex. This integrated palaeoenvironmental framework provides critical context for interpreting Late Pleistocene human adaptation and innovation in Europe.
Disciplines :
Archaeology Earth sciences & physical geography
Author, co-author :
Cartapanis, Olivier ; Aix-Marseille University, CNRS, IRD, Collège de France, INRAE, CEREGE, UMR7330, Aix-en-Provence, France
Bard, Edouard; Aix-Marseille University, CNRS, IRD, Collège de France, INRAE, CEREGE, UMR7330, Aix-en-Provence, France
Leroy, Suzanne A.G. ; Aix Marseille University, CNRS, Ministère de la Culture, LAMPEA, UMR 7269, Aix-en-Provence, France ; School of Environmental Sciences, University of Liverpool, Liverpool, United Kingdom
Chevalier, Manuel ; Institute of Geosciences, Sect. Meteorology, Rheinische Friedrich-Wilhelms-Universität Bonn, Bonn, Germany
Flas, Damien ; Université de Liège - ULiège > Département des sciences historiques > Archéologie préhistorique ; Aix Marseille University, CNRS, Ministère de la Culture, LAMPEA, UMR 7269, Aix-en-Provence, France
Devièse, Thibaut ; Aix-Marseille University, CNRS, IRD, Collège de France, INRAE, CEREGE, UMR7330, Aix-en-Provence, France
Language :
English
Title :
Regional climatic dynamics and cultural divergence in glacial western Europe
ANR - French National Research Agency European Union. Marie Skłodowska-Curie Actions
Funding text :
OC was supported by ANR NEHOS (PI: T. Devi\u00E8se, ANR-22-CE27-0016) and ANR MARCARA (PI: E. Bard, ANR-21-CE01-0023). The data collection benefited from a database developed within CYCLOCARB MSCA (PI: O. Cartapanis, MSCA No 897046). MC was supported by the German Federal Ministry of Education and Research ( BMBF ) as a Research for Sustainability initiative (FONA; https://www.fona.de/en ) through the PalMod Phase III project (grant no. 01LP2308B ). We are grateful to S. Badino, J. S. Carri\u00F3n, P. Stojakowits, C. Mayr, and S. Britzius who directly contributed their data. We thank Editor I. Hendy and one anonymous reviewer for their constructive feedback and insightful comments, which greatly improved the clarity and quality of this manuscript.
Allen, J.R.M., Brandt, U., Brauer, A., Hubberten, H.-W., Huntley, B., Keller, J., Kraml, M., et al. Rapid environmental changes in southern Europe during the last glacial period. Nature 400 (1999), 740–743.
Andersen, K.K., Svensson, A., Johnsen, S.J., Rasmussen, S.O., Bigler, M., Röthlisberger, R., Ruth, U., et al. The Greenland Ice Core Chronology 2005, 15–42ka. Part 1: constructing the time scale. Critical Quaternary Stratigraphy 25 (2006), 3246–3257.
Badino, F., Pini, R., Ravazzi, C., Chytrý, M., Bertuletti, P., Bortolini, E., Dudová, L., et al. High-resolution ecosystem changes pacing the millennial climate variability at the Middle to Upper Palaeolithic transition in NE-Italy. Sci. Rep., 13, 2023, 12478.
Banks, W.E., d'Errico, F., Zilhão, J., Human–climate interaction during the early upper paleolithic: testing the hypothesis of an adaptive shift between the proto-aurignacian and the early aurignacian. J. Hum. Evol. 64 (2013), 39–55.
Banks, W.E., Vignoles, A., Lacarrière, J., Morala, A., Klaric, L., A hierarchical bayesian examination of the chronological relationship between the noaillian and rayssian phases of the French middle gravettian. Quaternary, 7, 2024.
Bard, E., Arnold, M., Maurice, P., Duprat, J., Moyes, J., Duplessy, J.-C., Retreat velocity of the North Atlantic polar front during the last deglaciation determined by 14C accelerator mass spectrometry. Nature 328 (1987), 791–794.
Bard, E., Rostek, F., Jean Louis, T., Gendreau, S., Hydrological impact of Heinrich events in the subtropical northeast atlantic. Science (New York, N.Y.) 289 (2000), 1321–1324.
de Beaulieu, J.-L., Reille, M., The pollen sequence of les échets (France): a new element for the chronology of the upper Pleistocene. Géogr. Phys. Quaternaire 38 (1984), 3–9.
Bereiter, B., Eggleston, S., Schmitt, J., Nehrbass-Ahles, C., Stocker, T.F., Fischer, H., Kipfstuhl, S., Chappellaz, J., Revision of the EPICA Dome C CO2 record from 800 to 600 kyr before present. Geophys. Res. Lett. 42 (2015), 542–549.
Berger, A.L., LONG-TERM variations of caloric insolation resulting from earths orbital elements. Quat. Res. 9 (1978), 139–167.
Berger, A.L., The astronomical theory of paleoclimates. Berger, A., (eds.) Climatic Variations and Variability: Facts and Theories: NATO Advanced Study Institute First Course of the International School of Climatology, Ettore Majorana Center for Scientific Culture, 1981, Springer Netherlands, Dordrecht, Erice, Italy, 501–525 March 9–21, 1980.
Berger, A., Loutre, M.F., Insolation values for the climate of the last 10 million years. Quat. Sci. Rev. 10 (1991), 297–317.
Boemke, B., Maier, A., Schmidt, I., Römer, W., Lehmkuhl, F., Testing the representativity of Palaeolithic site distribution: the role of sampling bias in the european upper and Final Palaeolithic record. Quat. Sci. Rev., 316, 2023, 108220.
Bohm, E., Lippold, J., Gutjahr, M., Frank, M., Blaser, P., Antz, B., Fohlmeister, J., et al. Strong and deep Atlantic meridional overturning circulation during the last glacial cycle. Nature 517 (2015), 73–76.
Bond, G., Heinrich, H., Broecker, W., Labeyrie, L., McManus, J., Andrews, J., Huon, S., et al. Evidence for massive discharges of icebergs into the North Atlantic ocean during the last glacial period. Nature 360 (1992), 245–249.
Borzatti von Löwenstern, E., La grotta-riparo di Uluzzo C. Rivista di scienze preistoriche, 1965, 1–31.
ter Braak, C.J., van Dame, H., Inferring pH from diatoms: a comparison of old and new calibration methods. Hydrobiologia (The Hague) 178 (1989), 209–223.
Britzius, S., Dreher, F., Maisel, P., Sirocko, F., Vegetation patterns during the last 132,000 Years: a synthesis from twelve eifel maar sediment cores (Germany): the ELSA-23-pollen-stack. Quaternary, 7, 2024.
Brunck, H., Sirocko, F., Albert, J., The ELSA-Flood-Stack: a reconstruction from the laminated sediments of Eifel maar structures during the last 60000years. Global Planet. Change 142 (2016), 136–146.
Cartapanis, O., Jonkers, L., Moffa-Sanchez, P., Jaccard, S.L., de Vernal, A., Complex spatio-temporal structure of the Holocene thermal maximum. Nat. Commun., 13, 2022, 5662.
Charton, L., Combourieu-Nebout, N., Bertini, A., Lebreton, V., Peyron, O., Robles, M., Sassoon, D., Moncel, M.-H., Vegetation and climate changes during the Middle to Upper Palaeolithic transition in the southwestern Mediterranean: what happened to the last Neanderthals during Heinrich stadial 4?. Quat. Sci. Rev., 359, 2025, 109345.
Columbu, A., Chiarini, V., Spötl, C., Benazzi, S., Hellstrom, J., Cheng, H., De Waele, J., Speleothem record attests to stable environmental conditions during Neanderthal–modern human turnover in southern Italy. Nature Ecology & Evolution 4 (2020), 1188–1195.
Conte, M.H., Sicre, M.-A., Rã¼hlemann, C., Weber, J.C., Schulte, S., Schulz-Bull, D., Blanz, T., Global temperature calibration of the alkenone unsaturation index (UKâ€237) in surface waters and comparison with surface sediments. Geochem. Geophys. Geosystems, 7, 2006.
Davtian, N., Bard, E., A new view on abrupt climate changes and the bipolar seesaw based on paleotemperatures from Iberian Margin sediments. Proceedings of the National Academy of Sciences, vol.120, 2023, e2209558120.
Djindjian, F., Kozlowski, J., Otte, M., Le paléolithique supérieur en Europe. A. Colin, 1999.
Dommenget, D., The Ocean's role in continental climate variability and change. J. Clim. 22 (2009), 4939–4952.
d'Errico, F., Sánchez Goñi, M.F., Neandertal extinction and the millennial scale climatic variability of OIS 3. Quat. Sci. Rev. 22 (2003), 769–788.
d'Errico, F., Sanchez Goñi, M., Vanharen, M., L'impact de la variabilité climatique rapide des OIS3-2 sur le peuplement de l'Europe, 2006 [CrossRef Partial][Check multiple vol, first-page, last-page, msc3].
Eynaud, F., de Abreu, L., Voelker, A., Schönfeld, J., Salgueiro, E., Turon, J.-L., Penaud, A., et al. Position of the Polar Front along the western Iberian margin during key cold episodes of the last 45 ka. Geochem. Geophys. Geosystems, 10, 2009 n/a-n/a.
Favre, E., Escarguel, G., Suc, J.-P., Vidal, G., Thévenod, L., A contribution to deciphering the meaning of AP/NAP with respect to vegetation cover. Rev. Palaeobot. Palynol. 148 (2008), 13–35.
Felden, J., Möller, L., Schindler, U., Huber, R., Schumacher, S., Koppe, R., Diepenbroek, M., Glöckner, F.O., PANGAEA - data publisher for Earth & environmental science. Sci. Data, 10, 2023, 347.
Fletcher, W.J., Sánchez Goñi, M.F., Orbital- and sub-orbital-scale climate impacts on vegetation of the western Mediterranean basin over the last 48,000 yr. Quat. Res. 70 (2008), 451–464.
Fletcher, W.J., Sánchez Goñi, M.F., Allen, J.R.M., Cheddadi, R., Combourieu-Nebout, N., Huntley, B., Lawson, I., et al. Millennial-scale variability during the last glacial in vegetation records from Europe. Vegetation Response to Millennial-scale Variability during the Last Glacial 29 (2010), 2839–2864.
Fourcade, T., Goñi, M.F.S., Lahaye, C., Rossignol, L., Philippe, A., Environmental changes in SW France during the Middle to Upper Paleolithic transition from the pollen analysis of an eastern North Atlantic deep-sea core. Quat. Res. 110 (2022), 147–164.
Fourcade, T., Lahaye, C., Goñi, M.F.S., Cultural changes and adaptations to climatic and environmental changes of the last Neanderthals in southern France, Changements culturels et adaptations aux changements climatiques et environnementaux des derniers Néandertaliens dans le sud de la France. Université Michel de Montaigne - Bordeaux, III, 2022.
Ganopolski, A., Rahmstorf, S., Rapid changes of glacial climate simulated in a coupled climate model. Nature 409 (2001), 153–158.
Gowan, E.J., Zhang, X., Khosravi, S., Rovere, A., Stocchi, P., Hughes, A.L.C., Gyllencreutz, R., et al. A new global ice sheet reconstruction for the past 80 000 years. Nat. Commun., 12, 2021, 1199.
Guiot, J., de Beaulieu, J.L., Cheddadi, R., David, F., Ponel, P., Reille, M., The climate in Western Europe during the last Glacial/Interglacial cycle derived from pollen and insect remains. 200 KA of global change 103 (1993), 73–93.
Heaton, T.J., Bard, E., Bronk Ramsey, C., Butzin, M., Hatté, C., Hughen, K.A., Köhler, P., Reimer, P.J., A response to community questions on the MARINE20 radiocarbon age calibration curve: marine reservoir ages and the calibration of 14C samples from the oceans. Radiocarbon 65 (2023), 247–273.
Heaton, T.J., Butzin, M., Bard, E., Bronk Ramsey, C., Hughen, K.A., Köhler, P., Reimer, P.J., Marine radiocarbon calibration in polar regions: a simple approximate approach using MARINE20. Radiocarbon, 2023, 1–28.
Heinrich, H., Origin and consequences of cyclic ice rafting in the Northeast Atlantic Ocean during the past 130,000 years. Quat. Res. 29 (1988), 142–152.
Hemming, S.R., Heinrich events: massive late Pleistocene detritus layers of the North Atlantic and their global climate imprint. Rev. Geophys., 42, 2004.
Henry, L.G., McManus, J.F., Curry, W.B., Roberts, N.L., Piotrowski, A.M., Keigwin, L.D., North Atlantic ocean circulation and abrupt climate change during the last glaciation. Science 353 (2016), 470–474.
Heusser, L., Balsam, W.L., Pollen distribution in the northeast pacific ocean. Quat. Res. 7 (1977), 45–62.
Higham, T., Douka, K., Wood, R., Ramsey, C.B., Brock, F., Basell, L., Camps, M., et al. The timing and spatiotemporal patterning of Neanderthal disappearance. Nature 512 (2014), 306–309.
Higham, T., Frouin, M., Douka, K., Ronchitelli, A., Boscato, P., Benazzi, S., Crezzini, J., et al. Chronometric data and stratigraphic evidence support discontinuity between Neanderthals and early Homo sapiens in the Italian Peninsula. Nat. Commun., 15, 2024, 8016.
Hill, H.W., Flower, B.P., Quinn, T.M., Hollander, D.J., Guilderson, T.P., Laurentide Ice Sheet meltwater and abrupt climate change during the last glaciation. Paleoceanography, 21, 2006.
Höbig, N., Weber, M.E., Kehl, M., Weniger, G.-C., Julià, R., Melles, M., Fülöp, R.-H., et al. Lake Banyoles (northeastern Spain): a Last Glacial to Holocene multi-proxy study with regard to environmental variability and human occupation. Quat. Int. 274 (2012), 205–218.
Hublin, J.-J., The modern human colonization of western Eurasia: when and where?. Synchronising Environmental and Archaeological Records using Volcanic Ash Isochrons 118 (2015), 194–210.
Hussain, S.T., Floss, H., The role of river courses in organizing the cultural space of the Upper Paleolithic: examples from the Rhine, Rhône, Danube and Garonne. Modes de contacts et de déplacements au Paléolithique eurasiatique, 2014, 307–320 ERAUL 140, ArchéoLogique 5, Liège, Luxembourg.
Huybers, P., Eisenman, I., Integrated summer insolation calculations. NOAA/NCDC Paleoclimatology Program data contribution, 79, 2006.
Kageyama, M., Merkel, U., Otto-Bliesner, B., Prange, M., Abe-Ouchi, A., Lohmann, G., Ohgaito, R., et al. Climatic impacts of fresh water hosing under Last Glacial Maximum conditions: a multi-model study. Clim. Past 9 (2013), 935–953.
Kandel, A.W., Sommer, C., Kanaeva, Z., Bolus, M., Bruch, A.A., Groth, C., Haidle, M.N., et al. The ROCEEH Out of Africa Database (ROAD): a large-scale research database serves as an indispensable tool for human evolutionary studies. PLoS One, 18, 2023, e0289513.
Kappenberg, A., Amelung, W., Conze, N., Sirocko, F., Lehndorff, E., Fire–vegetation relationships during the last glacial cycle in a low mountain range (Eifel, Germany). Palaeogeogr. Palaeoclimatol. Palaeoecol., 562, 2021, 110140.
Kapsch, M.-L., Graversen, R.G., Economou, T., Tjernström, M., The importance of spring atmospheric conditions for predictions of the Arctic summer sea ice extent. Geophys. Res. Lett. 41 (2014), 5288–5296.
Kim, J.-H., Schouten, S., Hopmans, E.C., Donner, B., Sinninghe Damsté, J.S., Global sediment core-top calibration of the TEX86 paleothermometer in the ocean. Geochem. Cosmochim. Acta 72 (2008), 1154–1173.
Klein, K., Weniger, G.-C., Ludwig, P., Stepanek, C., Zhang, X., Wegener, C., Shao, Y., Assessing climatic impact on transition from Neanderthal to anatomically modern human population on Iberian Peninsula: a macroscopic perspective. Science Bulletin, 2023.
Klein, R., Archeology and the evolution of human behavior. Evol. Anthropol. 9 (2000), 17–36.
Koreneva, E.V., Marine palynological researches in the U.S.S.R. Marine Palynology 4 (1966), 565–574.
van Kreveld, S., Sarnthein, M., Erlenkeuser, H., Grootes, P., Jung, S., Nadeau, M.J., Pflaumann, U., Voelker, A., Potential links between surging ice sheets, circulation changes, and the Dansgaard-Oeschger Cycles in the Irminger Sea, 60–18 Kyr. Paleoceanography 15 (2000), 425–442.
Kutzbach, J., Gallimore, R., Harrison, S., Behling, P., Selin, R., Laarif, F., Climate and biome simulations for the past 21,000 years. Quat. Sci. Rev. 17 (1998), 473–506.
Laepple, T., Lohmann, G., Seasonal cycle as template for climate variability on astronomical timescales. Paleoceanography, 24, 2009.
Leroy, S.A.G., Arpe, K., Mikolajewicz, U., Vegetation context and climatic limits of the Early Pleistocene hominin dispersal in Europe. Early Human Evolution in the Western Palaearctic: Ecological Scenarios 30 (2011), 1448–1463.
Leroy, S.A.G., Giralt, S., Francus, P., Seret, G., The high sensitivity of the palynological record in the Vico maar lacustrine sequence (Latium, Italy) highlights the climatic gradient through Europe for the last 90 ka. Quat. Sci. Rev. 15 (1996), 189–201.
Li, C., Postl, A.K., Böhmer, T., Cao, X., Dolman, A.M., Herzschuh, U., Harmonized chronologies of a global late Quaternary pollen dataset (LegacyAge 1.0). Earth Syst. Sci. Data 14 (2022), 1331–1343.
Lippold, J., Grützner, J., Winter, D., Lahaye, Y., Mangini, A., Christl, M., Does sedimentary 231Pa/230Th from the Bermuda rise monitor past atlantic meridional overturning circulation?. Geophys. Res. Lett., 36, 2009.
Lisiecki, L.E., Raymo, M.E., A Pliocene-Pleistocene stack of 57 globally distributed benthic delta O-18 records. Paleoceanography, 20, 2005.
Liu, Z., Otto-Bliesner, B.L., He, F., Brady, E.C., Tomas, R., Clark, P.U., Carlson, A.E., et al. Transient simulation of last deglaciation with a new mechanism for bølling-allerød warming. Science, 325, 2009, 310.
Lougheed, B.C., Obrochta, S.P., A rapid, deterministic age-depth modeling routine for geological sequences with inherent depth uncertainty. Paleoceanogr. Paleoclimatol. 34 (2019), 122–133.
Lougheed, B., Obrochta, S., MatCal: open source bayesian 14C age calibration in matlab. J. Open Res. Software, 4, 2016.
Loutre, M.-F., Paillard, D., Vimeux, F., Cortijo, E., Does mean annual insolation have the potential to change the climate?. Earth Planet Sci. Lett. 221 (2004), 1–14.
Ludwig, P., Shao, Y., Kehl, M., Weniger, G.-C., The Last Glacial Maximum and Heinrich event I on the Iberian Peninsula: a regional climate modelling study for understanding human settlement patterns. Global Planet. Change 140 (2018), 34–47.
Luetscher, M., Boch, R., Sodemann, H., Spötl, C., Cheng, H., Edwards, R.L., Frisia, S., et al. North atlantic storm track changes during the last glacial maximum recorded by alpine speleothems. Nat. Commun., 6, 2015, 6344.
Marciani, G., Carmignani, L., Djakovic, I., Roussel, M., Arrighi, S., Rossini, M., Boschin, F., et al. The uluzzian and châtelperronian: No technological affinity in a shared chronological framework. Journal of Paleolithic Archaeology, 8, 2025, 3.
McManus, J.F., Francois, R., Gherardi, J.M., Keigwin, L.D., Brown-Leger, S., Collapse and rapid resumption of Atlantic meridional circulation linked to deglacial climate changes. Nature 428 (2004), 834–837.
Menviel, L., Timmermann, A., Friedrich, T., England, M.H., Hindcasting the continuum of Dansgaard–Oeschger variability: mechanisms, patterns and timing. Clim. Past 10 (2014), 63–77.
Monegato, G., Scardia, G., Hajdas, I., Rizzini, F., Piccin, A., The Alpine LGM in the boreal ice-sheets game. Sci. Rep., 7, 2017, 2078.
Moseley, G., Spötl, C., Brandstätter, S., Erhardt, T., Luetscher, M., Edwards, R., NALPS19: sub-orbital-scale climate variability recorded in northern Alpine speleothems during the last glacial period. Clim. Past 16 (2020), 29–50.
Movius, Jr H.L., The Châtelperronian in French archaeology: the evidence of Arcy-sur-Cure. Antiquity 43 (1969), 111–123.
Müller, U., Pross, J., Tzedakis, C., Gamble, C., Kotthoff, U., Schmiedl, G., Wulf, S., Christanis, K., The role of climate in the spread of modern humans into Europe. Quat. Sci. Rev. 30 (2011), 273–279.
Naughton, F., Sanchez Goñi, M.F., Desprat, S., Turon, J.-L., Duprat, J., Malaizé, B., Joli, C., et al. Present-day and past (last 25000 years) marine pollen signal off western Iberia. Mar. Micropaleontol. 62 (2007), 91–114.
Obreht, I., Hambach, U., Veres, D., Zeeden, C., Bösken, J., Stevens, T., Marković, S.B., et al. Shift of large-scale atmospheric systems over Europe during late MIS 3 and implications for Modern Human dispersal. Sci. Rep., 7, 2017, 5848.
Obreht, I., Hambach, U., Veres, D., Zeeden, C., Bösken, J., Stevens, T., Marković, S.B., et al. Shift of large-scale atmospheric systems over Europe during late MIS 3 and implications for Modern Human dispersal. Sci. Rep., 7, 2017, 5848.
Pedro, J.B., Andersson, C., Vettoretti, G., Voelker, A.H.L., Waelbroeck, C., Dokken, T.M., Jensen, M.F., et al. Dansgaard-Oeschger and Heinrich event temperature anomalies in the North Atlantic set by sea ice, frontal position and thermocline structure. Quat. Sci. Rev., 289, 2022, 107599.
Pini, R., Furlanetto, G., Vallé, F., Badino, F., Wick, L., Anselmetti, F.S., Bertuletti, P., et al. Linking North Atlantic and Alpine Last Glacial Maximum climates via a high-resolution pollen-based subarctic forest steppe record. Quat. Sci. Rev., 294, 2022, 107759.
Prentice, I.C., Cramer, W., Harrison, S.P., Leemans, R., Monserud, R.A., Solomon, A.M., Special paper: a global biome model based on plant physiology and dominance, soil properties and climate. J. Biogeogr. 19 (1992), 117–134.
Rasmussen, T.L., Thomsen, E., Moros, M., North Atlantic warming during Dansgaard-Oeschger events synchronous with Antarctic warming and out-of-phase with Greenland climate. Sci. Rep., 6, 2016, 20535.
Rathmann, H., Vizzari, M.T., Beier, J., Bailey, S.E., Ghirotto, S., Harvati, K., Human population dynamics in Upper Paleolithic Europe inferred from fossil dental phenotypes. Sci. Adv., 10, 2024, eadn8129.
Reimer, P.J., Austin, W.E.N., Bard, E., Bayliss, A., Blackwell, P.G., Bronk Ramsey, C., Butzin, M., et al. The IntCal20 northern hemisphere radiocarbon age calibration curve (0–55 cal kBP). Radiocarbon 62 (2020), 725–757.
Reimer, P.J., Reimer, R.W., A marine reservoir correction database and on-line interface. Radiocarbon 43 (2001), 461–463.
Roebroeks, W., Terra incognita: the Palaeolithic record of northwest Europe and the information potential of the southern North Sea. Netherlands Journal of Geosciences - Geologie en Mijnbouw 93 (2014), 43–53.
Rohling, E.J., Grant, K., Bolshaw, M., Roberts, A.P., Siddall, M., Hemleben, Ch, Kucera, M., Antarctic temperature and global sea level closely coupled over the past five glacial cycles. Nat. Geosci. 2 (2009), 500–504.
Rohling, E.J., Grant, K., Hemleben, C., Kucera, M., Roberts, A.P., Schmeltzer, I., Schulz, H., et al. New constraints on the timing of sea level fluctuations during early to middle marine isotope stage 3. Paleoceanography, 23, 2008.
Sanchez Goñi, M., Desprat, S., Fletcher, W., Morales-Molino, C., Naughton, F., Oliveira, D., Urrego, D., Zorzi, C., Pollen from the deep-sea: a breakthrough in the mystery of the ice ages. Front. Plant Sci., 9, 2018.
Sánchez Goñi, M.F., Regional impacts of climate change and its relevance to human evolution. Evolutionary Human Sciences, 2, 2020, e55.
Sánchez Goñi, M.F., Desprat, S., Daniau, A.-L., Bassinot, F.C., Polanco-Martínez, J.M., Harrison, S.P., Allen, J.R.M., et al. The ACER pollen and charcoal database: a global resource to document vegetation and fire response to abrupt climate changes during the last glacial period. Earth Syst. Sci. Data 9 (2017), 679–695.
Sánchez Goñi, M.F., Eynaud, F., Turon, J.L., Shackleton, N.J., High resolution palynological record off the Iberian margin: direct land-sea correlation for the Last Interglacial complex. Earth Planet Sci. Lett. 171 (1999), 123–137.
Sánchez Goñi, M.F., Landais, A., Fletcher, W.J., Naughton, F., Desprat, S., Duprat, J., Contrasting impacts of Dansgaard–Oeschger events over a western European latitudinal transect modulated by orbital parameters. Quat. Sci. Rev. 27 (2008), 1136–1151.
Schmidt, I., Gehlen, B., Winkler, K., Arrizabalaga, A., Arts, N., Bicho, N., Crombé, P., et al. Demographic responses to climatic changes during the final palaeolithic in Europe. bioRxiv: 2024.09.11.612499, 2024.
Shackleton, N.J., Hall, M.A., Vincent, E., Phase relationships between millennial-scale events 64,000–24,000 years ago. Paleoceanography 15 (2000), 565–569.
Shao, Y., Limberg, H., Klein, K., Wegener, C., Schmidt, I., Weniger, G.-C., Hense, A., Rostami, M., Human-existence probability of the Aurignacian techno-complex under extreme climate conditions. Quat. Sci. Rev., 263, 2021, 106995.
Shao, Y., Wegener, C., Klein, K., Schmidt, I., Weniger, G.-C., Reconstruction of human dispersal during Aurignacian on pan-European scale. Nat. Commun., 15, 2024, 7406.
Silvestrini, S., Romandini, M., Marciani, G., Arrighi, S., Carrera, L., Fiorini, A., López-García, J., et al. Integrated multidisciplinary ecological analysis from the Uluzzian settlement at the Uluzzo C Rock Shelter, south‐eastern Italy. J. Quat. Sci., 37, 2021.
Sirocko, F., Knapp, H., Dreher, F., Förster, M.W., Albert, J., Brunck, H., Veres, D., et al. The ELSA-vegetation-stack: reconstruction of landscape evolution zones (LEZ) from laminated eifel maar sediments of the last 60,000years. Global Planet. Change 142 (2016), 108–135.
Stanley, E.A., The application of palynology to oceanology with reference to the northwestern Atlantic. Deep Sea Res. Oceanogr. Abstr. 13 (1966), 921–939.
Stewart, J.R., Stringer, C.B., Human evolution out of Africa: the role of refugia and climate change. Science 335 (2012), 1317–1321.
Stott, L., Timmermann, A., Thunell, R., Southern hemisphere and deep-sea warming led deglacial atmospheric CO2 rise and tropical warming. Science 318 (2007), 435–438.
Sung, M.-K., An, S.-I., Shin, J., Park, J.-H., Yang, Y.-M., Kim, H.-J., Chang, M., Ocean fronts as decadal thermostats modulating continental warming hiatus. Nat. Commun., 14, 2023, 7777.
Svensson, A., Andersen, K.K., Bigler, M., Clausen, H.B., Dahl-Jensen, D., Davies, S.M., Johnsen, S.J., et al. A 60 000 year Greenland stratigraphic ice core chronology. Clim. Past 4 (2008), 47–57.
Svensson, A., Andersen, K.K., Bigler, M., Clausen, H.B., Dahl-Jensen, D., Davies, S.M., Johnsen, S.J., et al. The Greenland Ice Core Chronology 2005, 15–42ka. Part 2: comparison to other records. Critical Quaternary Stratigraphy 25 (2006), 3258–3267.
Tierney, J.E., Tingley, M.P., A Bayesian, spatially-varying calibration model for the TEX86 proxy. Geochem. Cosmochim. Acta 127 (2014), 83–106.
Tierney, J.E., Tingley, M.P., BAYSPLINE: a new calibration for the alkenone paleothermometer. Paleoceanogr. Paleoclimatol. 33 (2018), 281–301.
Timmermann, A., Raia, P., Mondanaro, A., Zollikofer, C.P.E., Ponce de León, M., Zeller, E., Yun, K.-S., Past climate change effects on human evolution. Nat. Rev. Earth Environ. 5 (2024), 701–716.
Tomasso, A., Porraz, G., Hunter-gatherers’ mobility and embedded raw material procurement strategies: a critical view from the Mediterranean Upper Paleolithic. Evol. Anthropol. Issues News Rev. 25 (2016), 164–174.
Veres, D., Bazin, L., Landais, A., Toyé Mahamadou Kele, H., Lemieux-Dudon, B., Parrenin, F., Martinerie, P., et al. The Antarctic ice core chronology (AICC2012): an optimized multi-parameter and multi-site dating approach for the last 120 thousand years. Clim. Past 9 (2013), 1733–1748.
Waelbroeck, C., Lougheed, B.C., Vazquez Riveiros, N., Missiaen, L., Pedro, J., Dokken, T., Hajdas, I., et al. Consistently dated Atlantic sediment cores over the last 40 thousand years. Sci. Data, 6, 2019, 165.
Wegwerth, A., Ganopolski, A., Ménot, G., Kaiser, J., Dellwig, O., Bard, E., Lamy, F., Arz, H., Black Sea temperature response to glacial millennial-scale climate variability. Geophys. Res. Lett., 42, 2015, 2015GL065499.
Williams, J.W., Grimm, E.C., Blois, J.L., Charles, D.F., Davis, E.B., Goring, S.J., Graham, R.W., et al. The Neotoma Paleoecology Database, a multiproxy, international, community-curated data resource. Quat. Res. 89 (2018), 156–177.
Wolff, E.W., Chappellaz, J., Blunier, T., Rasmussen, S., Svensson, A., Millennial-scale variability during the last glacial: the ice core record. Quat. Sci. Rev., 29, 2010.
Xu, G., Broadman, E., Dorado-Liñán, I., Klippel, L., Meko, M., Büntgen, U., De Mil, T., et al. Jet stream controls on European climate and agriculture since 1300 ce. Nature, 2024.
Zheng, P., Pedro, J.B., Jochum, M., Rasmussen, S.O., Lai, Z., Different trends in antarctic temperature and atmospheric CO2 during the last glacial. Geophys. Res. Lett., 48, 2021, e2021GL093868.
Zilhão, J., d'Errico, F., Banks, W.E., Teyssandier, N., A data‐driven paradigm shift for the middle‐to‐upper palaeolithic transition and the neandertal debate. Quaternary Environments and Humans, 2, 2024, 100037.